diff --git a/gerbil-manager-web/src/genetics/__tests__/genetics.test.ts b/gerbil-manager-web/src/genetics/__tests__/genetics.test.ts
index 9da7ca6..2a2aa22 100644
--- a/gerbil-manager-web/src/genetics/__tests__/genetics.test.ts
+++ b/gerbil-manager-web/src/genetics/__tests__/genetics.test.ts
@@ -222,8 +222,9 @@ describe('Farbschlag catalog', () => {
     expect(CATALOG).toHaveLength(CATALOG_SIZE)
     expect(CATALOG[0]).toMatchObject({ name: 'Pink Eyed White (PEW)', sortOrder: 0 })
     // Every row has a non-empty canonical genotype display string and unique name.
+    // GEN-3h: bracket notation (e[f], c[chm], c[h]) allowed in tokens.
     expect(new Set(CATALOG.map((c) => c.name)).size).toBe(CATALOG.length)
-    expect(CATALOG.every((c) => /^[A-Za-z?]+( [A-Za-z?]+){7}$/.test(c.canonicalGenotype))).toBe(true)
+    expect(CATALOG.every((c) => /^[A-Za-z[\]?-]+( [A-Za-z[\]?-]+){7}$/.test(c.canonicalGenotype))).toBe(true)
   })
 
   it('genotypeToFarbschlag (DATA-1 denormalization contract) returns the plain name', () => {
@@ -523,3 +524,99 @@ describe('GEN-3g: "-Hell" in variety name == cchm/ch het; hom == cchm/cchm', ()
     expect(name('aa cchmch DD EE gg PP spsp rere')).toBe('Zobel-Hell')
   })
 })
+
+describe('GEN-3h: breeder bracket-notation display + E-locus e-before-ef order', () => {
+  // ── Display symbols ────────────────────────────────────────────────────
+  it('ef displays as e[f], cchm as c[chm], ch as c[h]', () => {
+    // Fuchsschimmel: E=[ef,ef] hom
+    expect(toDisplayString(fromDisplayString('AA CC DD efef GG PP spsp rere'))).toBe(
+      'AA CC DD e[f]e[f] GG PP spsp rere',
+    )
+    // C-locus het: cchm + ch
+    expect(toDisplayString(fromDisplayString('aa cchmch DD EE GG PP spsp rere'))).toBe(
+      'aa c[chm]c[h] DD EE GG PP spsp rere',
+    )
+    // C-locus hom cchm
+    expect(toDisplayString(fromDisplayString('aa cchmcchm DD EE GG PP spsp rere'))).toBe(
+      'aa c[chm]c[chm] DD EE GG PP spsp rere',
+    )
+  })
+
+  // ── E-locus display order: E > e > e[f] ─────────────────────────────
+  it('Fuchsschimmel het pair {ef,e} displays as ee[f] (e before e[f])', () => {
+    // Stored canonical: [ef, e] (ef dominant over e in storage).
+    // Display must swap to [e, ef] per breeder convention.
+    const g = fromDisplayString('aa CC DD eef Gg Pp spsp rere')
+    expect(g.E).toEqual(['ef', 'e']) // storage order unchanged
+    expect(toDisplayString(g)).toBe('aa CC DD ee[f] Gg Pp spsp rere')
+  })
+
+  it('E+e stays Ee (E dominant over e, no swap needed)', () => {
+    expect(toDisplayString(fromDisplayString('aa CC DD Ee GG PP spsp rere'))).toBe(
+      'aa CC DD Ee GG PP spsp rere',
+    )
+  })
+
+  it('E+ef displays Ee[f] (E dominant stays first, ef renders as e[f])', () => {
+    expect(toDisplayString(fromDisplayString('aa CC DD Eef GG PP spsp rere'))).toBe(
+      'aa CC DD Ee[f] GG PP spsp rere',
+    )
+  })
+
+  // ── Julian oracle fixtures (HUMANQUESTION D3/D4) ─────────────────────
+  it('Tier C: oracle display string round-trips exactly (aa C- D- ee[f] Gg Pp spsp rere)', () => {
+    const display = 'aa C- D- ee[f] Gg Pp spsp rere'
+    const g = fromDisplayString(display)
+    expect(g.E).toEqual(['ef', 'e'])
+    expect(g.C).toEqual(['C', '?'])
+    expect(toDisplayString(g)).toBe(display)
+    // Farbschlag scope is outside GEN-3h; god confirmed colour is correct as-is.
+  })
+
+  it('Zuleika oracle: aa c[chm]c[h] DD Ee Gg PP spsp rere', () => {
+    const display = 'aa c[chm]c[h] DD Ee Gg PP spsp rere'
+    const g = fromDisplayString(display)
+    expect(g.C).toEqual(['cchm', 'ch'])
+    expect(g.E).toEqual(['E', 'e'])
+    expect(toDisplayString(g)).toBe(display)
+  })
+
+  it('Milka oracle: aa Cc[h] dd EE Gg P- Spsp rere', () => {
+    const display = 'aa Cc[h] dd EE Gg P- Spsp rere'
+    const g = fromDisplayString(display)
+    expect(g.C).toEqual(['C', 'ch'])
+    expect(g.D).toEqual(['d', 'd'])
+    expect(toDisplayString(g)).toBe(display)
+  })
+
+  // ── Parser accepts both forms ─────────────────────────────────────────
+  it('bracket input round-trips identically to internal-symbol input', () => {
+    expect(toDisplayString(fromDisplayString('AA c[chm]c[chm] DD EE GG PP spsp rere'))).toBe(
+      toDisplayString(fromDisplayString('AA cchmcchm DD EE GG PP spsp rere')),
+    )
+    expect(toDisplayString(fromDisplayString('AA CC DD e[f]e[f] GG PP spsp rere'))).toBe(
+      toDisplayString(fromDisplayString('AA CC DD efef GG PP spsp rere')),
+    )
+    expect(toDisplayString(fromDisplayString('AA CC DD ee[f] GG PP spsp rere'))).toBe(
+      toDisplayString(fromDisplayString('AA CC DD eef GG PP spsp rere')),
+    )
+  })
+
+  it('e[-] standalone: parses as [e,?], displays e-', () => {
+    const g = fromDisplayString('aa CC DD e[-] GG PP spsp rere')
+    expect(g.E).toEqual(['e', '?'])
+    expect(toDisplayString(g)).toBe('aa CC DD e- GG PP spsp rere')
+  })
+
+  it('CR-1a: Silvain oracle ee[-] parses without crash → [e,?], displays e-', () => {
+    // Real herdbook notation: ee[-] = fox allele e + unknown e-type second allele.
+    // The lookbehind rule strips the second e[-] → '?', leaving 'e?' for splitToken.
+    const input = 'aa c[chm]c[chm] Dd ee[-] Gg Pp Spsp'
+    const g = fromDisplayString(input)
+    expect(g.E).toEqual(['e', '?'])
+    expect(g.C).toEqual(['cchm', 'cchm'])
+    expect(g.D).toEqual(['D', 'd'])
+    expect(g.Sp).toEqual(['Sp', 'sp'])
+    expect(toDisplayString(g)).toBe('aa c[chm]c[chm] Dd e- Gg Pp Spsp rere')
+  })
+})
diff --git a/gerbil-manager-web/src/genetics/colorVarietySeed.generated.json b/gerbil-manager-web/src/genetics/colorVarietySeed.generated.json
index c696cbc..c6e634f 100644
--- a/gerbil-manager-web/src/genetics/colorVarietySeed.generated.json
+++ b/gerbil-manager-web/src/genetics/colorVarietySeed.generated.json
@@ -2,35 +2,35 @@
   {
     "name": "Pink Eyed White (PEW)",
     "english": "Pink Eyed White",
-    "canonicalGenotype": "AA chch DD EE GG pp spsp rere",
+    "canonicalGenotype": "AA c[h]c[h] DD EE GG pp spsp rere",
     "sortOrder": 0,
     "image": "rotaugen-weiss-pew-d-sep-e-sep.jpg"
   },
   {
     "name": "Hermelin",
     "english": "Dark Tailed White",
-    "canonicalGenotype": "aa chch DD EE GG PP spsp rere",
+    "canonicalGenotype": "aa c[h]c[h] DD EE GG PP spsp rere",
     "sortOrder": 1,
     "image": "hermelin.jpeg"
   },
   {
     "name": "Himalaya",
     "english": "Himalayan",
-    "canonicalGenotype": "AA chch DD EE GG PP spsp rere",
+    "canonicalGenotype": "AA c[h]c[h] DD EE GG PP spsp rere",
     "sortOrder": 2,
     "image": "himalaya.jpg"
   },
   {
     "name": "Zobel",
     "english": "Sable",
-    "canonicalGenotype": "aa cchmcchm DD EE gg PP spsp rere",
+    "canonicalGenotype": "aa c[chm]c[chm] DD EE gg PP spsp rere",
     "sortOrder": 3,
     "image": "zobel.jpeg"
   },
   {
     "name": "Rotaugenschimmel",
     "english": "Red-Eyed Roan",
-    "canonicalGenotype": "AA CC DD efef GG pp spsp rere",
+    "canonicalGenotype": "AA CC DD e[f]e[f] GG pp spsp rere",
     "sortOrder": 4,
     "image": "rotaugen-schimmel.jpg"
   },
@@ -168,7 +168,7 @@
   },
   {
     "name": "Orangeschimmel",
-    "canonicalGenotype": "AA CC DD efef GG PP spsp rere",
+    "canonicalGenotype": "AA CC DD e[f]e[f] GG PP spsp rere",
     "sortOrder": 25,
     "image": "schimmel-orangeschimmel.jpg"
   },
@@ -210,31 +210,31 @@
   },
   {
     "name": "Silberschimmel",
-    "canonicalGenotype": "AA CC DD efef gg PP spsp rere",
+    "canonicalGenotype": "AA CC DD e[f]e[f] gg PP spsp rere",
     "sortOrder": 32,
     "image": "silberschimmel.jpg"
   },
   {
     "name": "Polarfuchsschimmel",
-    "canonicalGenotype": "AA CC DD efef gg PP spsp rere",
+    "canonicalGenotype": "AA CC DD e[f]e[f] gg PP spsp rere",
     "sortOrder": 33,
     "image": "polarfuchsschimmel.jpg"
   },
   {
     "name": "Algierfuchsschimmel",
-    "canonicalGenotype": "AA CC DD efef GG PP spsp rere",
+    "canonicalGenotype": "AA CC DD e[f]e[f] GG PP spsp rere",
     "sortOrder": 34,
     "image": "algierfuchsschimmel.jpg"
   },
   {
     "name": "Kohlfuchsschimmel",
-    "canonicalGenotype": "aa CC DD efef GG PP spsp rere",
+    "canonicalGenotype": "aa CC DD e[f]e[f] GG PP spsp rere",
     "sortOrder": 35,
     "image": "kohlfuchsschimmel.jpg"
   },
   {
     "name": "Blaufuchsschimmel",
-    "canonicalGenotype": "aa CC DD efef gg PP spsp rere",
+    "canonicalGenotype": "aa CC DD e[f]e[f] gg PP spsp rere",
     "sortOrder": 36,
     "image": "blaufuchsschimmel.jpg"
   },
@@ -252,7 +252,7 @@
   },
   {
     "name": "Goldfuchsschimmel",
-    "canonicalGenotype": "AA CC DD efef GG pp spsp rere",
+    "canonicalGenotype": "AA CC DD e[f]e[f] GG pp spsp rere",
     "sortOrder": 39,
     "image": "goldfuchsschimmel.jpg"
   },
@@ -270,7 +270,7 @@
   },
   {
     "name": "Rotfuchsschimmel",
-    "canonicalGenotype": "aa CC DD efef GG pp spsp rere",
+    "canonicalGenotype": "aa CC DD e[f]e[f] GG pp spsp rere",
     "sortOrder": 42,
     "image": "rotfuchsschimmel.jpg"
   },
@@ -282,7 +282,7 @@
   },
   {
     "name": "Kohlfuchsschimmel, hell",
-    "canonicalGenotype": "aa CC DD efef GG PP spsp rere",
+    "canonicalGenotype": "aa CC DD e[f]e[f] GG PP spsp rere",
     "sortOrder": 44,
     "image": "kohlfuchsschimmel-hell.jpg"
   },
@@ -318,89 +318,89 @@
   },
   {
     "name": "Marder",
-    "canonicalGenotype": "aa cchmcchm DD EE GG PP spsp rere",
+    "canonicalGenotype": "aa c[chm]c[chm] DD EE GG PP spsp rere",
     "sortOrder": 50,
     "image": "marder.JPG"
   },
   {
     "name": "Siam",
-    "canonicalGenotype": "aa cchmch DD EE GG PP spsp rere",
+    "canonicalGenotype": "aa c[chm]c[h] DD EE GG PP spsp rere",
     "sortOrder": 51,
     "image": "siam-marder-hell.JPG"
   },
   {
     "name": "Zobel-Hell",
-    "canonicalGenotype": "aa cchmch DD EE gg PP spsp rere",
+    "canonicalGenotype": "aa c[chm]c[h] DD EE gg PP spsp rere",
     "sortOrder": 52,
     "image": "zobel-hell.jpg"
   },
   {
     "name": "CP-Agouti",
-    "canonicalGenotype": "AA cchmcchm DD EE GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[chm] DD EE GG PP spsp rere",
     "sortOrder": 53,
     "image": "agouti-cp.jpg"
   },
   {
     "name": "CP-Agouti-Hell",
-    "canonicalGenotype": "AA cchmch DD EE GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[h] DD EE GG PP spsp rere",
     "sortOrder": 54
   },
   {
     "name": "CP-Silberagouti",
-    "canonicalGenotype": "AA cchmcchm DD EE gg PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[chm] DD EE gg PP spsp rere",
     "sortOrder": 55,
     "image": "silberagouti-cp.JPG"
   },
   {
     "name": "CP-Silberagouti-Hell",
-    "canonicalGenotype": "AA cchmch DD EE gg PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[h] DD EE gg PP spsp rere",
     "sortOrder": 56
   },
   {
     "name": "CP-Algierfuchs",
-    "canonicalGenotype": "AA cchmcchm DD ee GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[chm] DD ee GG PP spsp rere",
     "sortOrder": 57,
     "image": "algierfuchs-cp.jpg"
   },
   {
     "name": "CP-Algierfuchs-Hell",
-    "canonicalGenotype": "AA cchmch DD ee GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[h] DD ee GG PP spsp rere",
     "sortOrder": 58
   },
   {
     "name": "CP-Polarfuchs",
-    "canonicalGenotype": "AA cchmcchm DD ee gg PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[chm] DD ee gg PP spsp rere",
     "sortOrder": 59,
     "image": "polarfuchs-cp.jpg"
   },
   {
     "name": "CP-Polarfuchs-Hell",
-    "canonicalGenotype": "AA cchmch DD ee gg PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[h] DD ee gg PP spsp rere",
     "sortOrder": 60
   },
   {
     "name": "CP-Fuchs",
-    "canonicalGenotype": "AA cchmcchm dd ee GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[chm] dd ee GG PP spsp rere",
     "sortOrder": 61
   },
   {
     "name": "CP-Fuchs-Hell",
-    "canonicalGenotype": "AA cchmch dd ee GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[h] dd ee GG PP spsp rere",
     "sortOrder": 62
   },
   {
     "name": "CP-Blaufuchs",
-    "canonicalGenotype": "AA cchmcchm dd ee gg PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[chm] dd ee gg PP spsp rere",
     "sortOrder": 63
   },
   {
     "name": "CP-Orangeschimmel",
-    "canonicalGenotype": "AA cchmcchm DD efef GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[chm] DD e[f]e[f] GG PP spsp rere",
     "sortOrder": 64
   },
   {
     "name": "CP-Orangeschimmel-Hell",
-    "canonicalGenotype": "AA cchmch DD efef GG PP spsp rere",
+    "canonicalGenotype": "AA c[chm]c[h] DD e[f]e[f] GG PP spsp rere",
     "sortOrder": 65
   }
 ]
diff --git a/gerbil-manager-web/src/genetics/genotype.ts b/gerbil-manager-web/src/genetics/genotype.ts
index 6441c93..5a54606 100644
--- a/gerbil-manager-web/src/genetics/genotype.ts
+++ b/gerbil-manager-web/src/genetics/genotype.ts
@@ -74,6 +74,32 @@ export function wildType(): Genotype {
   return out
 }
 
+/**
+ * GEN-3h: breeder bracket-notation display symbols.
+ * STORAGE symbols (ef / cchm / ch) are frozen; only the rendered form changes.
+ */
+const DISPLAY_SYMBOL: Readonly<Partial<Record<string, string>>> = {
+  ef: 'e[f]',
+  cchm: 'c[chm]',
+  ch: 'c[h]',
+}
+function displaySymbol(allele: string): string {
+  return DISPLAY_SYMBOL[allele] ?? allele
+}
+
+/**
+ * GEN-3h: E-locus display order — breeder convention is E > e > e[f].
+ * Storage/dominance order is E > ef > e; display swaps ef and e so that
+ * a Fuchsschimmel (E=[ef,e] stored) renders as "ee[f]" not "e[f]e".
+ */
+const E_DISPLAY_RANK: Readonly<Record<string, number>> = { E: 0, e: 1, ef: 2 }
+
+function displayPair(locus: LocusKey, pair: AllelePair): AllelePair {
+  if (locus !== 'E') return pair
+  const rank = (x: string) => E_DISPLAY_RANK[x] ?? Number.MAX_SAFE_INTEGER
+  return rank(pair[0]) <= rank(pair[1]) ? pair : [pair[1], pair[0]]
+}
+
 /**
  * Compact display string, e.g. "Aa CC Dd EE GG Pp spsp rere".
  * The Sls locus is OMITTED when wild-type (sl/sl) so legacy 8-locus strings and
@@ -81,12 +107,18 @@ export function wildType(): Genotype {
  * (e.g. "… spsp rere Slsl"). Round-trips: a missing Sls re-parses to sl/sl.
  * GEN-3c: unknown alleles are STORED as '?' but DISPLAYED as '-' (breeder
  * convention) — e.g. ['C','?'] renders "C-".
+ * GEN-3h: sub-alleles use breeder bracket notation (e[f], c[chm], c[h]);
+ * E-locus display order is E > e > e[f] (e before e[f] in het pairs).
  */
 export function toDisplayString(g: Genotype): string {
   return LOCUS_ORDER.filter(
     (locus) => locus !== 'Sls' || !(g.Sls[0] === 'sl' && g.Sls[1] === 'sl'),
   )
-    .map((locus) => (g[locus][0] + g[locus][1]).replace(/\?/g, '-'))
+    .map((locus) => {
+      const [a, b] = displayPair(locus, g[locus])
+      return displaySymbol(a) + displaySymbol(b)
+    })
+    .map((s) => s.replace(/\?/g, '-'))
     .join(' ')
 }
 
@@ -146,8 +178,19 @@ function normalizeToken(tok: string): string | null {
   if (t === 'WP') t = 'Slsl'
   t = t.replace(/S\(l\)/g, 'Sl').replace(/s\(l\)/g, 'sl')
   t = t.replace(/Uw/g, 'G').replace(/uw/g, 'g')
-  // GEN-3c: '-' is the breeder's UNKNOWN marker on input; store internally as '?'
-  // (the frozen storage contract keeps '?'; only DISPLAY renders '-').
+  // GEN-3h: accept bracket display notation → canonical internal symbols.
+  t = t.replace(/e\[f\]/g, 'ef')     // Schimmel allele display form → internal
+  t = t.replace(/c\[chm\]/g, 'cchm') // Colourpoint display form → internal
+  t = t.replace(/c\[h\]/g, 'ch')     // Himalayan display form → internal
+  // CR-1a: allele-prefixed bracket-unknown like ee[-] (Silvain).
+  // When e[-] is PRECEDED by a letter it is the second unknown allele in a
+  // 2-allele token (e.g. ee[-] → e + e[-] → e + ?).  Lookbehind strips only
+  // the e[-] part; the leading allele stays. Standalone e[-] falls through to
+  // the generic [-]→? rule below (which makes the bracket-dash a wildcard,
+  // leaving the leading allele intact for splitToken).
+  t = t.replace(/(?<=[A-Za-z])e\[-\]/g, '?')
+  t = t.replace(/\[-\]/g, '?')       // bare/standalone bracket-unknown → wildcard
+  // GEN-3c: plain dash is the breeder's UNKNOWN marker on input; store internally as '?'.
   t = t.replace(/-/g, '?')
   return t
 }