GEN-3c: unknown allele displays as '-' (storage stays '?'); parser accepts '-' and skips fully-unknown tokens; de.ts hints

gerbil-manager-web/src/genetics/genotype.ts

@@ -79,12 +79,14 @@ export function wildType(): Genotype {
  * The Sls locus is OMITTED when wild-type (sl/sl) so legacy 8-locus strings and
  * the colour catalog stay byte-identical; it only appears for WP/Sls carriers
  * (e.g. "… spsp rere Slsl"). Round-trips: a missing Sls re-parses to sl/sl.
+ * GEN-3c: unknown alleles are STORED as '?' but DISPLAYED as '-' (breeder
+ * convention) — e.g. ['C','?'] renders "C-".
  */
 export function toDisplayString(g: Genotype): string {
   return LOCUS_ORDER.filter(
     (locus) => locus !== 'Sls' || !(g.Sls[0] === 'sl' && g.Sls[1] === 'sl'),
   )
-    .map((locus) => g[locus][0] + g[locus][1])
+    .map((locus) => (g[locus][0] + g[locus][1]).replace(/\?/g, '-'))
     .join(' ')
 }
 
@@ -144,6 +146,9 @@ function normalizeToken(tok: string): string | null {
   if (t === 'WP') t = 'Slsl'
   t = t.replace(/S\(l\)/g, 'Sl').replace(/s\(l\)/g, 'sl')
   t = t.replace(/Uw/g, 'G').replace(/uw/g, 'g')
+  // GEN-3c: '-' is the breeder's UNKNOWN marker on input; store internally as '?'
+  // (the frozen storage contract keeps '?'; only DISPLAY renders '-').
+  t = t.replace(/-/g, '?')
   return t
 }
 
@@ -193,7 +198,9 @@ export function fromDisplayString(input: string): Genotype {
     const [a, b] = splitToken(token)
     const refAllele = a === WILDCARD ? b : a
     if (refAllele === WILDCARD) {
-      throw new Error(`Token "${token}" is fully unknown; cannot infer its locus`)
+      // Fully-unknown token ("--"/"??", e.g. a positional placeholder): the locus
+      // can't be inferred — skip it (defaults to wild-type), don't throw.
+      continue
     }
     const locus = ALLELE_TO_LOCUS[refAllele]
     if (!locus) throw new Error(`Unknown allele "${refAllele}" in token "${token}"`)